Research Professor in the Department of Integrative Ecology at the Estación Biológica de Doñana, CSIC.
His research focuses on the ecological and evolutionary consequences of mutualistic interactions between animals and plants, exploring how coevolution shapes megadiverse communities where such mutualisms play an essential role. He divides his fieldwork time between the Spanish Mediterranean forest and the Atlantic forest of Brazil.
A former coordinator of the Biodiversity, Ecology and Global Change program of the Spanish National Research Plan and chair of the ERC Starting Grants panel in Ecology, Evolution and Environmental Biology, he holds the Mercer Award of the Ecological Society of America, and the Rey Jaime I Prize for Environmental Conservation. He also holds an honorary professorship at the University of Seville, and serves on the editorial board of the Annual Review of Ecology, Evolution, and Systematics.